{"id":57,"date":"2016-09-19T16:04:17","date_gmt":"2016-09-19T23:04:17","guid":{"rendered":"https:\/\/sites.chapman.edu\/fudge\/?page_id=57"},"modified":"2016-09-19T16:04:17","modified_gmt":"2016-09-19T23:04:17","slug":"publications","status":"publish","type":"page","link":"https:\/\/sites.chapman.edu\/fudge\/publications\/","title":{"rendered":"Publications"},"content":{"rendered":"

[et_pb_section fb_built=”1″ fullwidth=”on” _builder_version=”4.16″ global_colors_info=”{}”][et_pb_fullwidth_header title=”Publications” text_orientation=”center” _builder_version=”4.16″ background_color=”rgba(255, 255, 255, 0)” background_image=”https:\/\/sites.chapman.edu\/fudge\/files\/2016\/08\/Fudge-banner-1-172du2o.jpg” background_url=”https:\/\/sites.chapman.edu\/fudge\/files\/2016\/08\/Fudge-banner-1-172du2o.jpg” saved_tabs=”all” global_colors_info=”{}” button_one_text_size__hover_enabled=”off” button_two_text_size__hover_enabled=”off” button_one_text_color__hover_enabled=”off” button_two_text_color__hover_enabled=”off” button_one_border_width__hover_enabled=”off” button_two_border_width__hover_enabled=”off” button_one_border_color__hover_enabled=”off” button_two_border_color__hover_enabled=”off” button_one_border_radius__hover_enabled=”off” button_two_border_radius__hover_enabled=”off” button_one_letter_spacing__hover_enabled=”off” button_two_letter_spacing__hover_enabled=”off” button_one_bg_color__hover_enabled=”off” button_two_bg_color__hover_enabled=”off”][\/et_pb_fullwidth_header][\/et_pb_section][et_pb_section fb_built=”1″ admin_label=”section” _builder_version=”4.16″ global_colors_info=”{}”][et_pb_row padding_mobile=”off” column_padding_mobile=”on” admin_label=”row” _builder_version=”4.19.5″ background_size=”initial” background_position=”top_left” background_repeat=”repeat” custom_margin=”|auto|-42px|auto||” custom_padding=”3px|||1px||” global_colors_info=”{}”][et_pb_column type=”4_4″ _builder_version=”4.16″ custom_padding=”|||” global_colors_info=”{}” custom_padding__hover=”|||”][et_pb_text _builder_version=”4.25.2″ background_size=”initial” background_position=”top_left” background_repeat=”repeat” custom_padding=”2px|||||” use_border_color=”off” global_colors_info=”{}”]<\/p>\n

PDF \u00a0 Hossain, M.T., Eom, W., Layak, P., Kim, J., Darling, C., Lowe, A., Fudge, D.S., Tawfick, S.H., & Ewoldt, R.H. (2025). Design Principles for Deployable Fibers Inspired by Hagfish Defense. Advanced Science<\/em>, e12414.<\/span><\/span><\/a>\u00a0<\/span><\/span><\/p>\n

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DOI <\/span>\u00a0 <\/a>Hossain, M.T., Piorkowski, D.<\/span>, Lowe, A., Wonsik, E., Shetty, A., Tawfick, S.H., Fudge, D.S., \u00a0Ewoldt, R.H. (2025). Physics of unravelling and micromechanics of hagfish threads<\/span>. Journal of Royal Society Interface<\/em>, 22(229)<\/span><\/span><\/p>\n

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DOI \u00a0 <\/a>Hossain, M.T., Eom, W., Shah, A.<\/span>, Lowe, A., Fudge, D.S., Tawfick, S.H., Ewoldt, R.H. (2025). The critical plastocapillary number for a Newtonian liquid filament embedded into a viscoplastic fluid<\/span>. Journal of Non-Newtonian Fluid Mechanics<\/em>, 343, 105440<\/span><\/span><\/p>\n

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DOI \u00a0 <\/a><\/span>Eom, W., Hossain, M.T., Parasramka, V., Kim, J.,\u00a0<\/span>Siu, R.W.Y., Sanders, K.A., Piorkowski, D., Lowe, A., Koh, H., De Volder, M.F.L., Fudge, D.S.,<\/span> Ewoldt, R.H., Tawfick, S.H. (2025). Fast 3D printing of fine, continuous, and soft fibers via embedded solvent exchange<\/span>. Nature communications<\/em>, 16<\/span>(842)<\/span><\/p>\n

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PDF \u00a0 <\/span><\/a>Hossain, M.T., Piorkowski, D., Lowe, A., Eom, W., Shetty, A., Tawfick, S.H., Fudge, D.S., and Ewoldt, R.H.<\/span> (2025). Physics of unravelling and micromechanics of hagfish threads<\/span>. Journal of the Royal Society Interface<\/span><\/i>, <\/span>22<\/i>(229), p.20250503.\u00a0<\/span><\/span><\/p>\n

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PDF<\/a> \u00a0 <\/span>Fudge, D.S., Lee, J., Guillen, K., Donatelli, C.M., Lowe, A., Arnold, L., Kahale-Lua, K., Quinteros, C., Ly, P., Atkins, L. and Bressman, N. (2024). Biphasic burrowing in Atlantic hagfish (Myxine limosa<\/em>). Journal of Experimental Biology<\/i>, 227<\/i>(12), jeb247544.\u00a0<\/span><\/p>\n

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PDF \u00a0 <\/a><\/span>Zeng, Y., Plachetzki, D.C., Nieders, K., Campbell, H., Cartee, M., Pankey, M.S., Guillen, K. and Fudge, D., (2023). Epidermal threads reveal the origin of hagfish slime. Elife<\/i>, 12<\/i>, p.e81405.<\/span><\/p>\n

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PDF \u00a0 <\/a><\/span>Taylor, L., Chaudhary, G., Jain, G., Lowe, A., Hupe, A., Negishi, A., Zeng, Y., Ewoldt, R.H. and Fudge, D.S.(2023). Mechanisms of gill-clogging by hagfish slime. Journal of the Royal Society Interface<\/i>, 20<\/i>(200), 20220774.<\/span><\/p>\n

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PDF <\/a><\/span>\u00a0 \u00a0\u00a0Zeng Y., Petrichko S, Nieders K., Plachetzki D, and Fudge D.S. (2021) Evolution of a remarkable intracellular polymer and extreme cell allometry in hagfishes. Current Biology<\/i>, 31: 5062-8.<\/span><\/p>\n

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PDF <\/a><\/span>\u00a0 \u00a0 \u00a0Bressman, N., & Fudge<\/span>, D.S. (2021). From reductionionism to synthesis: the case of hagfish slime. Comparative Biochemistry and Physiology, Part B, 255,\u00a0<\/i>1-10.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Mincarone, M. M., Plachetzki, D., McCord, C.L., Winegard, T.M., Fernholm, B, Gonzalez, C.J., & Fudge, D.S. (2021). Review of the hagfishes (Myxinidae<\/em>) from the Galapagos Islands, with descriptions of four new species and their phylogenetic relationships. Zoological Journal of the Linnean Society, 192,<\/i> 453-474.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 McCord, C., Whiteley, E., Liang, J., Trejo, C., Caputo, R., Itehua, E., Hasan, H., Hernandez, S., Jagnandan, K., & Fudge, D. S. (2020). Concentration effects of three common fish anesthetics on Pacific hagfish (Eptatretus stoutii<\/em>). Fish Physiol Biochem<\/i>, 46<\/i>, 931-944.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., Ferraro, F.N., Siwiecki, S.A., Hupe, A., & Jain, G. (2020). A new model of hagfish slime mucous vesicle stabilization and deployment. Langmuir<\/i>, 36<\/i>(24), 6681-6689.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Black, K.L., Fudge, D. S., Jarvis, W.M.C., & Robinson, B.W. (2019). Functional plasticity in lamellar autotomy by larval damselflies in response to predatory larval dragonfly cues. Evolutionary Ecology<\/i>, 33<\/i>, 257-272.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Jain, G., Starksen, M., Singh, K., Hoang, C., Yancey, P., McCord, C., & Fudge, D. S. (2019). High concentrations of trimethylamines in slime glands inhibit skein unraveling in Pacific Hagfish. Journal of Experimental Biology <\/i>, 222<\/i>(22).<\/span><\/p>\n

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PDF<\/a>\u00a0 \u00a0Bernards Jr., M. A., Schorno, S., McKenzie, E., Winegard, T. M., Oke, I., Plachetzki, D. & Fudge, D. S. (2018). Unraveling inter-species differences in hagfish slime skein deployment<\/span>.\u00a0Journal of Experimental Biology, 221,\u00a0<\/em>1-11.<\/span><\/span><\/span><\/p>\n

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PDF<\/a>\u00a0<\/span> \u00a0Schorno, S., Gillis, T. E. & Fudge, D. S. (2018). Cellular mechanisms of slime gland refilling in Pacific hagfish (Eptatretus stoutii<\/em>).\u00a0Journal of Experimental Biology, 221,\u00a0<\/em>1-11.<\/span><\/span><\/span><\/p>\n

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PDF<\/a>\u00a0 \u00a0<\/span>Schorno, S., Gillis, T. E. & Fudge, D. S. (2018).\u00a0Emptying and refilling of slime glands in Atlantic (Myxine\u00a0<\/em><\/span>glutinosa<\/em>) and Pacific (Eptatretus stoutii<\/em>) hagfishes.\u00a0Journal of Experimental Biology, 221,\u00a0<\/em>1-8.<\/span><\/p>\n

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PDF<\/a><\/span>\u00a0 \u00a0 \u00a0Freedman, C.\u00a0 R. & Fudge, D. S. (2017). Hagfish Houdinis: biomechanics and behavior of squeezing through small openings. Journal of Experimental Biology,<\/em>\u00a0220<\/em>, 822-827.<\/span><\/p>\n

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PDF<\/a><\/span>\u00a0 \u00a0 \u00a0Boggett, S., Stiles, J-L, Summers, A. P. & Fudge, D. S. (2017). Flaccid skin protects hagfishes from shark bites. J. R. Soc.<\/em><\/span>
Interface,<\/em> 14: 20170765.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Weatherbee-Martin, N., Xu, L., Hupe, A., Kreplak, L., Fudge, D. S., Liu, X., & Rainey, J. K. (2016). Identification of wet-spinning and post-spin stretching methods amenable to recombinant spider aciniform silk. Biomacromolecules<\/i>, 17<\/i>(8), 2737-2746.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Icardo, J. M., Colvee, E., Schorno, S., Lauriano, E. R., Fudge, D. S., Glover, C. N., & Zaccone, G. (2016). Morphological analysis of the hagfish heart. II. The venous pole and the pericardium. Journal of Morphology<\/i>, 277<\/i>(7), 853-865.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., & Schorno, S. (2016). The hagfish gland thread cell: a fiber-producing cell involved in predator defense. \u00a0Cells<\/i>, 5<\/i>(2), 25.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., Schorno, S., & Ferraro, S. (2014). Physiology, biomechanics, and biomimetics of hagfish slime.\u00a0Annual Review of Biochemistry<\/i>, (0).<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Won, G. J., Fudge, D. S., & Choh, V. (2015). The effects of actomyosin disruptors on the mechanical integrity of the avian crystalline lens.\u00a0Molecular vision<\/i>,\u00a021<\/i>, 98.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Winegard, T., Herr, J., Mena, C., Lee, B., Dinov, I., Bird, D., Bernards, M Jr., Hobel, S., Valkenburgh, B., Toga, A., & Fudge, D. (2014). Coiling and maturation of a high-performance fibre in hagfish slime gland thread cells. Nature communications<\/i>,\u00a05<\/i>.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Pinto, N., Yang, F. C., Negishi, A., Rheinst\u00e4dter, M. C., Gillis, T. E., & Fudge, D. S. (2014). Self-assembly enhances the strength of fibers made from vimentin intermediate filament proteins.\u00a0Biomacromolecules<\/i>,\u00a015<\/i>(2), 574-581.<\/span><\/p>\n

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PDF <\/a> \u00a0\u00a0\u00a0Bernards, M. A., Oke, I., Heyland, A., & Fudge, D. S. (2014). Spontaneous unraveling of hagfish slime thread skeins is mediated by a seawater-soluble protein adhesive.\u00a0The Journal of experimental biology<\/i>,\u00a0217<\/i>(8), 1263-1268.<\/span><\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S. (2014). Fifty years of JR Platt\u2019s strong inference.\u00a0The Journal of experimental biology<\/i>,\u00a0217<\/i>(8), 1202-1204.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Herr, J. E., Clifford, A., Goss, G. G., & Fudge, D. S. (2014). Defensive slime formation in Pacific hagfish requires Ca2+ and aquaporin mediated swelling of released mucin vesicles.\u00a0The Journal of experimental biology<\/i>,\u00a0217: 2288-2296<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Gleason, J. E., Fudge, D. S., & Robinson, B. W. (2014). Eco-mechanics of lamellar autotomy in larval damselflies.\u00a0The Journal of experimental biology<\/i>, 217<\/i>(2), 185-191.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Negishi, A., Armstrong, C. L., Kreplak, L., Rheinstadter, M. C., Lim, L. T., Gillis, T. E., & Fudge, D. S. (2012). The production of fibers and films from solubilized hagfish slime thread proteins.\u00a0Biomacromolecules<\/i>,\u00a013<\/i>(11), 3475-3482.<\/span><\/p>\n

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PDF<\/a><\/span> \u00a0\u00a0\u00a0 Greenberg, D. A., & Fudge, D. S. (2013). Regulation of hard \u03b1-keratin mechanics via control of intermediate filament hydration: matrix squeeze revisited.\u00a0Proceedings of the Royal Society B: Biological Sciences<\/i>,\u00a0280<\/i>(1750), 20122158.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Beriault, D. R., Haddad, O., McCuaig, J. V., Robinson, Z. J., Russell, D., Lane, E. B., & Fudge, D. S. (2012). The mechanical behavior of mutant K14-R125P keratin bundles and networks in NEB-1 keratinocytes.\u00a0PloS one<\/i>,\u00a07<\/i>(2), e31320.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., McCuaig, J. V., Van Stralen, S., Hess, J. F., Wang, H., Mathias, R. T., & FitzGerald, P. G. (2011). Intermediate filaments regulate tissue size and stiffness in the murine lens.\u00a0Investigative ophthalmology & visual science<\/i>, 52<\/i>(6), 3860-3867.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0Ewoldt, R. H., Winegard, T. M., & Fudge, D. S. (2011). Non-linear viscoelasticity of hagfish slime.\u00a0International Journal of Non-Linear Mechanics<\/i>, 46<\/i>(4), 627-636.<\/span><\/p>\n

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PDF<\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., Hillis, S., Levy, N., & Gosline, J. M. (2010). Hagfish slime threads as a biomimetic model for high performance protein fibres.\u00a0Bioinspiration & biomimetics<\/i>,\u00a05<\/i>(3), 035002.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Szewciw, L. J., De Kerckhove, D. G., Grime, G. W., & Fudge, D. S. (2010). Calcification provides mechanical reinforcement to whale baleen \u03b1-keratin. Proceedings of the Royal Society B: Biological Sciences<\/i>, 277(1694), 2597-2605, rspb20100399.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Herr, J. E., Winegard, T. M., O\u2019Donnell, M. J., Yancey, P. H., & Fudge, D. S. (2010). Stabilization and swelling of hagfish slime mucin vesicles.\u00a0The Journal of experimental biology<\/i>,\u00a0213<\/i>(7), 1092-1099.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Winegard, T. M., & Fudge, D. S. (2010). Deployment of hagfish slime thread skeins requires the transmission of mixing forces via mucin strands.\u00a0The Journal of Experimental Biology<\/i>,\u00a0213<\/i>(8), 1235-1240.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 LeBlanc, D. M., Wood, C. M., Fudge, D. S., & Wright, P. A. (2010). A fish out of water: gill and skin remodeling promotes osmo-and ionoregulation in the mangrove killifish Kryptolebias marmoratus.\u00a0Physiological and Biochemical Zoology<\/i>,\u00a083<\/i>(6), 932-949.<\/span><\/p>\n

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PDF\u00a0<\/a>\u00a0\u00a0\u00a0<\/span>Fudge, D.S., Winegard, T., Ewoldt, R.H. Beriault, D., Szewciw, L., and McKinley, G.H. (2009) From ultra-soft slime to hard alpha-keratins: the many lives of intermediate filaments.\u00a0Integrative and\u00a0Comparative Biology<\/i>, 49<\/i>(1), 32-39.<\/span><\/p>\n

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PDF <\/a><\/span>\u00a0\u00a0\u00a0 Fudge, D. S., Szewciw, L. J., & Schwalb, A. N. (2009). Morphology and development of blue whale baleen: an annotated translation of Tycho Tullberg\u2019s classic 1883 paper.\u00a0Aquatic Mammals<\/i>,\u00a035<\/i>(2), 226.<\/span><\/p>\n

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PDF <\/a> \u00a0\u00a0\u00a0<\/span>Fudge, D., Russell, D., Beriault, D., Moore, W., Lane, E. B., & Vogl, A. W. (2008). The intermediate filament network in cultured human keratinocytes is remarkably extensible and resilient.\u00a0PLoS One<\/i>,\u00a03<\/i>(6), e2327.<\/span><\/p>\n

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PDF <\/a> \u00a0\u00a0\u00a0 <\/span>Kreplak, L., & Fudge, D. (2007). Biomechanical properties of intermediate filaments: from tissues to single filaments and back.\u00a0Bioessays<\/i>,\u00a029<\/i>(1), 26-35.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Lim, J., Fudge, D. S., Levy, N., & Gosline, J. M. (2006). Hagfish slime ecomechanics: testing the gill-clogging hypothesis.\u00a0Journal of experimental biology<\/i>,\u00a0209<\/i>(4), 702-710.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 John M. Gosline, Douglas S. Fudge, and Paul A. Guerette. (2006). Alpha-helical protein based materials and methods for making same (patent).<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., Levy, N., Chiu, S., & Gosline, J. M. (2005). Composition, morphology and mechanics of hagfish slime.\u00a0Journal of experimental biology<\/i>,208<\/i>(24), 4613-4625.<\/span><\/p>\n

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PDF <\/a><\/span>\u00a0\u00a0\u00a0 Fudge, D. S., & Gosline, J. M. (2004). Molecular design of the \u03b1\u2013keratin composite: insights from a matrix\u2013free model, hagfish slime threads.\u00a0Proceedings of the Royal Society of London. Series B: Biological Sciences<\/i>,271<\/i>(1536), 291-299.<\/span><\/p>\n

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PDF <\/a>\u00a0\u00a0\u00a0Fudge, D. S., Gardner, K. H., Forsyth, V. T., Riekel, C., & Gosline, J. M. (2003). The mechanical properties of hydrated intermediate filaments: insights from hagfish slime threads.\u00a0Biophysical Journal<\/i>,\u00a085<\/i>(3), 2015-2027.<\/span><\/span><\/p>\n

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PDF<\/a> \u00a0 <\/span>Fudge, D.S., Lee, J., Guillen, K., Donatelli, C.M., Lowe, A., Arnold, L., Kahale-Lua, K., Quinteros, C., Ly, P., Atkins, L. and Bressman, N. (2024). Biphasic burrowing in Atlantic hagfish (Myxine limosa). Journal of Experimental Biology<\/i>, 227<\/i>(12), jeb247544.\u00a0<\/span><\/span><\/p>\n

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PDF<\/a> \u00a0 <\/span>Fudge, D.S., Lee, J., Guillen, K., Donatelli, C.M., Lowe, A., Arnold, L., Kahale-Lua, K., Quinteros, C., Ly, P., Atkins, L. and Bressman, N. (2024). Biphasic burrowing in Atlantic hagfish (Myxine limosa). Journal of Experimental Biology<\/i>, 227<\/i>(12), jeb247544.\u00a0<\/span><\/p>\n

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PDF \u00a0 <\/a><\/span>Zeng, Y., Plachetzki, D.C., Nieders, K., Campbell, H., Cartee, M., Pankey, M.S., Guillen, K. and Fudge, D., (2023). Epidermal threads reveal the origin of hagfish slime. Elife<\/i>, 12<\/i>, p.e81405.<\/span><\/p>\n

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PDF \u00a0 <\/a><\/span>Taylor, L., Chaudhary, G., Jain, G., Lowe, A., Hupe, A., Negishi, A., Zeng, Y., Ewoldt, R.H. and Fudge, D.S.(2023). Mechanisms of gill-clogging by hagfish slime. Journal of the Royal Society Interface<\/i>, 20<\/i>(200), 20220774.<\/span><\/p>\n

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PDF <\/a><\/span>\u00a0 \u00a0\u00a0Zeng Y., Petrichko S, Nieders K., Plachetzki D, and Fudge D.S. (2021) Evolution of a remarkable intracellular polymer and extreme cell allometry in hagfishes. Current Biology<\/i>, 31: 5062-8.<\/span><\/p>\n

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PDF <\/a><\/span>\u00a0 \u00a0 \u00a0Bressman, N., & Fudge<\/span>, D.S. (2021). From reductionionism to synthesis: the case of hagfish slime. Comparative Biochemistry and Physiology, Part B, 255,\u00a0<\/i>1-10.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Mincarone, M. M., Plachetzki, D., McCord, C.L., Winegard, T.M., Fernholm, B, Gonzalez, C.J., & Fudge, D.S. (2021). Review of the hagfishes (Myxinidae<\/em>) from the Galapagos Islands, with descriptions of four new species and their phylogenetic relationships. Zoological Journal of the Linnean Society, 192,<\/i> 453-474.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 McCord, C., Whiteley, E., Liang, J., Trejo, C., Caputo, R., Itehua, E., Hasan, H., Hernandez, S., Jagnandan, K., & Fudge, D. S. (2020). Concentration effects of three common fish anesthetics on Pacific hagfish (Eptatretus stoutii<\/em>). Fish Physiol Biochem<\/i>, 46<\/i>, 931-944.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., Ferraro, F.N., Siwiecki, S.A., Hupe, A., & Jain, G. (2020). A new model of hagfish slime mucous vesicle stabilization and deployment. Langmuir<\/i>, 36<\/i>(24), 6681-6689.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Black, K.L., Fudge, D. S., Jarvis, W.M.C., & Robinson, B.W. (2019). Functional plasticity in lamellar autotomy by larval damselflies in response to predatory larval dragonfly cues. Evolutionary Ecology<\/i>, 33<\/i>, 257-272.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Jain, G., Starksen, M., Singh, K., Hoang, C., Yancey, P., McCord, C., & Fudge, D. S. (2019). High concentrations of trimethylamines in slime glands inhibit skein unraveling in Pacific Hagfish. Journal of Experimental Biology <\/i>, 222<\/i>(22).<\/span><\/p>\n

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PDF<\/a>\u00a0 \u00a0Bernards Jr., M. A., Schorno, S., McKenzie, E., Winegard, T. M., Oke, I., Plachetzki, D. & Fudge, D. S. (2018). Unraveling inter-species differences in hagfish slime skein deployment<\/span>.\u00a0Journal of Experimental Biology, 221,\u00a0<\/em>1-11.<\/span><\/span><\/span><\/p>\n

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PDF<\/a>\u00a0<\/span> \u00a0Schorno, S., Gillis, T. E. & Fudge, D. S. (2018). Cellular mechanisms of slime gland refilling in Pacific hagfish (Eptatretus stoutii<\/em>).\u00a0Journal of Experimental Biology, 221,\u00a0<\/em>1-11.<\/span><\/span><\/span><\/p>\n

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PDF<\/a>\u00a0 \u00a0<\/span>Schorno, S., Gillis, T. E. & Fudge, D. S. (2018).\u00a0Emptying and refilling of slime glands in Atlantic (Myxine\u00a0<\/em><\/span>glutinosa<\/em>) and Pacific (Eptatretus stoutii<\/em>) hagfishes.\u00a0Journal of Experimental Biology, 221,\u00a0<\/em>1-8.<\/span><\/p>\n

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PDF<\/a><\/span>\u00a0 \u00a0 \u00a0Freedman, C.\u00a0 R. & Fudge, D. S. (2017). Hagfish houdinis: biomechanics and behavior of squeezing through small openings.\u00a0Journal of Experimental Biology,<\/em>\u00a0220<\/em>, 822-827.<\/span><\/p>\n

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PDF<\/a><\/span>\u00a0 \u00a0 \u00a0Boggett, S., Stiles, J-L, Summers, A. P. & Fudge, D. S. (2017). Flaccid skin protects hagfishes from shark bites. J. R. Soc.<\/em><\/span>
Interface,<\/em> 14: 20170765.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Weatherbee-Martin, N., Xu, L., Hupe, A., Kreplak, L., Fudge, D. S., Liu, X., & Rainey, J. K. (2016). Identification of wet-spinning and post-spin stretching methods amenable to recombinant spider aciniform silk. Biomacromolecules<\/i>, 17<\/i>(8), 2737-2746.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Icardo, J. M., Colvee, E., Schorno, S., Lauriano, E. R., Fudge, D. S., Glover, C. N., & Zaccone, G. (2016). Morphological analysis of the hagfish heart. II. The venous pole and the pericardium. Journal of Morphology<\/i>, 277<\/i>(7), 853-865.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., & Schorno, S. (2016). The hagfish gland thread cell: a fiber-producing cell involved in predator defense. \u00a0Cells<\/i>, 5<\/i>(2), 25.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., Schorno, S., & Ferraro, S. (2014). Physiology, biomechanics, and biomimetics of hagfish slime.\u00a0Annual Review of Biochemistry<\/i>, (0).<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Won, G. J., Fudge, D. S., & Choh, V. (2015). The effects of actomyosin disruptors on the mechanical integrity of the avian crystalline lens.\u00a0Molecular vision<\/i>,\u00a021<\/i>, 98.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Winegard, T., Herr, J., Mena, C., Lee, B., Dinov, I., Bird, D., \u2026 & Fudge, D. (2014). Coiling and maturation of a high-performance fibre in hagfish slime gland thread cells.\u00a0Nature communications<\/i>,\u00a05<\/i>.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Pinto, N., Yang, F. C., Negishi, A., Rheinst\u00e4dter, M. C., Gillis, T. E., & Fudge, D. S. (2014). Self-assembly enhances the strength of fibers made from vimentin intermediate filament proteins.\u00a0Biomacromolecules<\/i>,\u00a015<\/i>(2), 574-581.<\/span><\/p>\n

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PDF <\/a> \u00a0\u00a0\u00a0Bernards, M. A., Oke, I., Heyland, A., & Fudge, D. S. (2014). Spontaneous unraveling of hagfish slime thread skeins is mediated by a seawater-soluble protein adhesive.\u00a0The Journal of experimental biology<\/i>,\u00a0217<\/i>(8), 1263-1268.<\/span><\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S. (2014). Fifty years of JR Platt\u2019s strong inference.\u00a0The Journal of experimental biology<\/i>,\u00a0217<\/i>(8), 1202-1204.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Herr, J. E., Clifford, A., Goss, G. G., & Fudge, D. S. (2014). Defensive slime formation in Pacific hagfish requires Ca2+ and aquaporin mediated swelling of released mucin vesicles.\u00a0The Journal of experimental biology<\/i>,\u00a0217: 2288-2296<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Gleason, J. E., Fudge, D. S., & Robinson, B. W. (2014). Eco-mechanics of lamellar autotomy in larval damselflies.\u00a0The Journal of experimental biology<\/i>, 217<\/i>(2), 185-191.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Negishi, A., Armstrong, C. L., Kreplak, L., Rheinstadter, M. C., Lim, L. T., Gillis, T. E., & Fudge, D. S. (2012). The production of fibers and films from solubilized hagfish slime thread proteins.\u00a0Biomacromolecules<\/i>,\u00a013<\/i>(11), 3475-3482.<\/span><\/p>\n

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PDF<\/a><\/span> \u00a0\u00a0\u00a0 Greenberg, D. A., & Fudge, D. S. (2013). Regulation of hard \u03b1-keratin mechanics via control of intermediate filament hydration: matrix squeeze revisited.\u00a0Proceedings of the Royal Society B: Biological Sciences<\/i>,\u00a0280<\/i>(1750), 20122158.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Beriault, D. R., Haddad, O., McCuaig, J. V., Robinson, Z. J., Russell, D., Lane, E. B., & Fudge, D. S. (2012). The mechanical behavior of mutant K14-R125P keratin bundles and networks in NEB-1 keratinocytes.\u00a0PloS one<\/i>,\u00a07<\/i>(2), e31320.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., McCuaig, J. V., Van Stralen, S., Hess, J. F., Wang, H., Mathias, R. T., & FitzGerald, P. G. (2011). Intermediate filaments regulate tissue size and stiffness in the murine lens.\u00a0Investigative ophthalmology & visual science<\/i>, 52<\/i>(6), 3860-3867.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0Ewoldt, R. H., Winegard, T. M., & Fudge, D. S. (2011). Non-linear viscoelasticity of hagfish slime.\u00a0International Journal of Non-Linear Mechanics<\/i>, 46<\/i>(4), 627-636.<\/span><\/p>\n

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PDF<\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., Hillis, S., Levy, N., & Gosline, J. M. (2010). Hagfish slime threads as a biomimetic model for high performance protein fibres.\u00a0Bioinspiration & biomimetics<\/i>,\u00a05<\/i>(3), 035002.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Szewciw, L. J., De Kerckhove, D. G., Grime, G. W., & Fudge, D. S. (2010). Calcification provides mechanical reinforcement to whale baleen \u03b1-keratin. Proceedings of the Royal Society B: Biological Sciences<\/i>, 277(1694), 2597-2605, rspb20100399.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Herr, J. E., Winegard, T. M., O\u2019Donnell, M. J., Yancey, P. H., & Fudge, D. S. (2010). Stabilization and swelling of hagfish slime mucin vesicles.\u00a0The Journal of experimental biology<\/i>,\u00a0213<\/i>(7), 1092-1099.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Winegard, T. M., & Fudge, D. S. (2010). Deployment of hagfish slime thread skeins requires the transmission of mixing forces via mucin strands.\u00a0The Journal of Experimental Biology<\/i>,\u00a0213<\/i>(8), 1235-1240.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 LeBlanc, D. M., Wood, C. M., Fudge, D. S., & Wright, P. A. (2010). A fish out of water: gill and skin remodeling promotes osmo-and ionoregulation in the mangrove killifish Kryptolebias marmoratus.\u00a0Physiological and Biochemical Zoology<\/i>,\u00a083<\/i>(6), 932-949.<\/span><\/p>\n

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PDF\u00a0<\/a>\u00a0\u00a0\u00a0<\/span>Fudge, D.S., Winegard, T., Ewoldt, R.H. Beriault, D., Szewciw, L., and McKinley, G.H. (2009) From ultra-soft slime to hard alpha-keratins: the many lives of intermediate filaments.\u00a0Integrative and\u00a0Comparative Biology<\/i>, 49<\/i>(1), 32-39.<\/span><\/p>\n

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PDF <\/a><\/span>\u00a0\u00a0\u00a0 Fudge, D. S., Szewciw, L. J., & Schwalb, A. N. (2009). Morphology and development of blue whale baleen: an annotated translation of Tycho Tullberg\u2019s classic 1883 paper.\u00a0Aquatic Mammals<\/i>,\u00a035<\/i>(2), 226.<\/span><\/p>\n

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PDF <\/a> \u00a0\u00a0\u00a0<\/span>Fudge, D., Russell, D., Beriault, D., Moore, W., Lane, E. B., & Vogl, A. W. (2008). The intermediate filament network in cultured human keratinocytes is remarkably extensible and resilient.\u00a0PLoS One<\/i>,\u00a03<\/i>(6), e2327.<\/span><\/p>\n

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PDF <\/a> \u00a0\u00a0\u00a0 <\/span>Kreplak, L., & Fudge, D. (2007). Biomechanical properties of intermediate filaments: from tissues to single filaments and back.\u00a0Bioessays<\/i>,\u00a029<\/i>(1), 26-35.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Lim, J., Fudge, D. S., Levy, N., & Gosline, J. M. (2006). Hagfish slime ecomechanics: testing the gill-clogging hypothesis.\u00a0Journal of experimental biology<\/i>,\u00a0209<\/i>(4), 702-710.<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 John M. Gosline, Douglas S. Fudge, and Paul A. Guerette. (2006). Alpha-helical protein based materials and methods for making same (patent).<\/span><\/p>\n

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PDF <\/a><\/span> \u00a0\u00a0\u00a0 Fudge, D. S., Levy, N., Chiu, S., & Gosline, J. M. (2005). Composition, morphology and mechanics of hagfish slime.\u00a0Journal of experimental biology<\/i>,208<\/i>(24), 4613-4625.<\/span><\/p>\n

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PDF <\/a><\/span>\u00a0\u00a0\u00a0 Fudge, D. S., & Gosline, J. M. (2004). Molecular design of the \u03b1\u2013keratin composite: insights from a matrix\u2013free model, hagfish slime threads.\u00a0Proceedings of the Royal Society of London. Series B: Biological Sciences<\/i>,271<\/i>(1536), 291-299.<\/span><\/p>\n

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PDF <\/a>\u00a0\u00a0\u00a0Fudge, D. S., Gardner, K. H., Forsyth, V. T., Riekel, C., & Gosline, J. M. (2003). The mechanical properties of hydrated intermediate filaments: insights from hagfish slime threads.\u00a0Biophysical Journal<\/i>,\u00a085<\/i>(3), 2015-2027.<\/span><\/span><\/p>\n

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PDF \u00a0 Hossain, M.T., Eom, W., Layak, P., Kim, J., Darling, C., Lowe, A., Fudge, D.S., Tawfick, S.H., & Ewoldt, R.H. (2025). Design Principles for Deployable Fibers Inspired by Hagfish Defense. Advanced Science, e12414.\u00a0   DOI \u00a0 Hossain, M.T., Piorkowski, D., Lowe, A., Wonsik, E., Shetty, A., Tawfick, S.H., Fudge, D.S., \u00a0Ewoldt, R.H. (2025). Physics […]<\/p>\n","protected":false},"author":83,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"_et_pb_use_builder":"on","_et_pb_old_content":"","_et_gb_content_width":"","footnotes":""},"class_list":["post-57","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/sites.chapman.edu\/fudge\/wp-json\/wp\/v2\/pages\/57","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/sites.chapman.edu\/fudge\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/sites.chapman.edu\/fudge\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/sites.chapman.edu\/fudge\/wp-json\/wp\/v2\/users\/83"}],"replies":[{"embeddable":true,"href":"https:\/\/sites.chapman.edu\/fudge\/wp-json\/wp\/v2\/comments?post=57"}],"version-history":[{"count":0,"href":"https:\/\/sites.chapman.edu\/fudge\/wp-json\/wp\/v2\/pages\/57\/revisions"}],"wp:attachment":[{"href":"https:\/\/sites.chapman.edu\/fudge\/wp-json\/wp\/v2\/media?parent=57"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}